***Pardon My Progress - Site Undergoing Metamorphosis***

My research interests are broad and encompass a diversity of taxa and ecosystems. However, many revolve around a central theme in understanding the impact of human activities, intentional and unintentional, on species and biotic communities. Through my research I hope to shed light on solutions to the negative anthropogenic effects we have on Nature. I hope to achieve this by adhering to the scientific method, employing ecological statistics and by developing a better understanding of species life histories and ecological communities. However, I also recognize that the majority of our environmental problems arise due to a conflict of value systems among cultures, societies, and individuals. To this end I recognize that science represents only one perspective in a multi-perspectival world and it alone will not provide a sustainable solution to the environmental crises we face. More recently I have employed the developing field of Integral Ecology to provide a framework to address the role that science should play in addressing our environmental problems, including endangered species recovery and habitat management.


Current Research Interests

Impacts of Herbicides on Endangered Butterflies

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Next to habitat loss and fragmentation, invasive plant species represent the second largest threat to several endangered butterfly species in the Pacific Northwest as well as other regions of the world. Control of invasive woody plants, forbs, and grasses has historically focused on the use of mowing, fire, and biological control to manage invasive plant species. However, these methods can have severe impacts on species occupying those systems. For example, fire can kill a large percentage of larvae and pupa butterfly in the path of the flames. For this reason land managers and conservation biologists working with endangered butterflies are unable to employ these methods at effective invasive weed management scales for fear of extirpating or severely impacting local butterfly populations. Increasingly land managers and conservation biologists have turned to herbicides as alternative management tool. Many herbicides have been developed to target specific taxa (e.g. grasses) while causing little to no visible impact on native non-target plant species. However, little research has been conducted to date on the potential impacts, both positive and negative, of herbicide exposure on butterflies. Through a combination of greenhouse and field studies I am investigating the impacts of grass-specific and broad-spectrum herbicides on butterfly behavior, morphology, survival, and fecundity as part of my dissertation research at Washington State University - Vancouver in the Conservation Biology lab of Dr. Cheryl Schultz.

Butterfly Friendly Wine

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Less than one percent of the native prairie habitat remains in the Willamette Valley of western Oregon. The once expansive prairies have now largely been lost to afforestation, invasion by shrubs and grasses, urban development, and conversion to agriculture. Viticulture is particularly prevalent and large vineyards can be found throughout the region. Many of these vineyard lie adjacent to or near prairie patches hosting endangered butterflies including the endangered Fender's Blue Butterfly. Working in collaboration with federal agencies and private landowners I am interested in developing a "Fender's Friendly" wine certification program. This program would aim to increase the biotic prairie value of wineries through planting of native nectar and host plant species, reducing pesticide use, and increasing public awareness of butterfly conservation in region.

Impacts of Tidal Flat Loss on the Avifauna
in the Yellow Sea Ecoregion


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The tidal flats in the Yellow Sea ecoregion are disappearing at an alarming rate. These rich ecosystems provide important stopovers for literally hundreds of thousands of migratory shorebirds as they migrate north to their breeding grounds on the Russian and Alaskan Arctic tundra and when they return southward in the fall. Exact measures of tidal flat loss are difficult to ascertain. However, a number of large scale tidal flat reclamation projects have been completed in the past decade including the environmentally devastating Saemangeum reclamation project that destroyed over 400 square kilometers of tidal flat and estuary. The Saemangeum estuary once hosted internationally important numbers of the Spoon-billed Sandpiper and Great Knot as well a host of other bird species. Remaining tidal flats still support a wide array of species including the globally vulnerable Saunder's Gull and Chinese Egret and provide feeding grounds for endangered Nordmann's Greenshank and Black-faced Spoonbill. I am working with the avian conservation group Birds Korea to quantify remaining tidal flats. I am using high-resolution publicly available satellite imagery (Daum and Google Earth) and Arc-GIS mapping technologies to relate remaining tidal flats to bird distributions and to focus conservation efforts in the region. (click here for more information)


Ecology and Impacts of Forestry Practices on
the Pacific Banana Slug

(I know they don't have wings)

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The Pacific Banana Slug is the second largest slug in the world and can be found in damp forests from California to southeast Alaska. While there is a vast amount of literature on slug ecology and impacts in an agriculture little is known about forest slug ecology. I am interested in learning more about the basic life history traits and ecology of Pacific banana slugs using recently developed fluorescent pigment marking for use in mark-recapture techniques to explore the impacts of forestry practices on the species.

Impacts of Wildlife Based Recreation on Wildlife

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Wildlife based recreation (e.g. photography, birding, ecotourism) is among the fastest growing forms of recreation in the United States. Recreation wildlife watchers can affect wildlife behavior in a variety of way. I am interested in quantifying the impacts of recreational activities associated with wildlife based recreation. I have been working with student Ryan Blazer and professor Dr. Pat Magee Western State College of Colorado to investigate the impacts of birders at a Gunnison Sage-Grouse lek in the Gunnison Basin of western Colorado. In addition, I am interested in determining the impacts of playback, a method employed by birders to lure birds into view using audio recording of the species, on nesting success and territory establishment and maintenance.

Using Data Generated by Amateur Ornithologists to
Document Changes in Phenology and Range of Birds

Birding is one of the most popular and rapidly growing recreational activities in the United States. Vast quantities of data are recorded by birders and stored in a variety of open digital sources including: submissions to rare bird records committees, breeding bird atlases, birding list-servs and newsletters, and recently through online checklist programs such as eBird. Using these sources I am interested in documenting range expansion and changes in migratory bird phenology using this data. In addition, I am interested in determining how accurately the birding community can detect species declines.

Past Research Interests

Sperm Morphometry and Ejaculate Characteristics
in Gunnison and Greater Sage-Grouse


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My Sr. thesis (2008) project at Western State College of Colorado focused on comparing sperm morphometry and natural ejaculate characteristics of the two species of Centrocercus grouse occurring in Colorado: Greater Sage-Grouse (Centrocercus nevadensis) and Gunnison Sage-Grouse (C. minimus). In spring 2008 we collected natural ejaculates from strutting male Greater Sage-grouse (Centrocercus urophasianus) in Moffat County, Colorado and from Gunnison Sage-grouse in Saguache County, Colorado. Natural ejaculates were collected by placing freeze-dried female greater sage-grouse in the copulatory position fitted with false cloacas on leks. We compared the ejaculate characteristics of sperm concentration, percent viable sperm, percent motile sperm, relative motility of sperm, percent structurally normal sperm, and the types and predominance of abnormal sperm in ejaculate samples. This study was intended as a preliminary investigation into the potential effects of inbreeding on Gunnison sage-grouse. With the exception of sperm concentration, we found few differences in ejaculate characteristics between the two species. However, we compared and found significant differences in the sperm morphometry of the two species. This study is the first to document natural ejaculate characteristics and sperm morphometry in a non-passerine bird species utilizing a polygnous lek mating system.


Life History and Conservation of the
Queen Alexandra's Birdwing Butterfly


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The Queen Alexandra's Birdwing Butterfly (QABB) (Ornithoptera alexandrae) is the largest butterfly in the world.The male's wingspan averages an impressive 178 mm and the larger females average an impressive 200 mm. However, females with wingspans of up to 250 mm (8+ inches) have been recorded. QABB is restricted to a small geographic area in the Oro (Northern) Province of mainland Papua New Guinea. Here they reside in the lush secondary and primary forests growing on volcanic soils. Throughout its range QABB occurs at low densities and an observer can spend several days in suitable habitat without encountering an adult. Three widely separated populations of QABB can be found in the region. One population can be found found on the volcanic Popondetta Plain but widespread logging and development of plantations, primarily oil palm, have destroyed and severely fragmented forest habitats in the region. A second high elevation population occurs in the Afore region and appears to be less impacted by anthropogenic activities. A third isolated population in the Cape Ward region has yet to be quantified in any systematic way. Due to the isolation and small size of each population, habitat loss, and demand for this species in the specimen trade the species has been listed as CITES I, endangered. During the summer of 2008 I worked for Conservation International and with the Oro provincial wildlife office to document the status of the species in the region, shed light on the species life history, and to identify and make contact with traditional land owners, with QABB residing on their land, interested in conserving the species.

Butterfly Communities and Abundance in
Mechanically Treated Sagebrush-Steppe

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Vocal Ecology of Boreal Owls

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Impacts of Grazing on Prairie Butterflies

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